Changes in hydrology, too much water, but especially too little moisture, represent ominous, proximate threats to the world’s insect fauna, as well as the plants upon which more than 50% of all insects feed. In aridlands and elsewhere, water stress has been identified as a primary driver of rapid insect population declines. In addition to direct losses through desiccation, droughts have detrimental knock-on effects that trigger further losses. While increased fire risk may be the most obvious of these, many other knock-on effects are driving declines: e.g., the lowering of water tables, diminished snowpack, changed soil hydrology, the collapse of soil microfaunas, diminishment of nectar resources, and the lowering of hostplant fitness over times of water stress.